1. Abdel-Baki, G. K., Shaddad, M. A. K., Mostafa, D., & Rafat, A. S. (2018). The effect of seed presoaking with KNO3 on seed germination, proline, protein pattern, ß-amylase and mineral composition of two faba bean cultivars treated with NaCl. Egyptian Journal of Botany, 58(3), 445-461. [
DOI:10.21608/ejbo.2018]
2. Abraham, J., & Florentine, S. (2021). Licorice (Glycyrrhiza glabra) extracts-suitable pharmacological interventions for COVID-19? A review. Plants, 10(12), 2600. [
DOI:10.3390/plants10122600] [
PMID] [
PMCID]
3. Afsharzadeh, N., Paltram, R., Jungwirth, A., Tabrizi, L., Nazeri, V., Kalantari, H., Halbwirth, H., Samiei, L., Sheehan, H., & Shokrpour, M. (2025). Advancing Glycyrrhiza glabra L. cultivation and hairy root transformation and elicitation for future metabolite overexpression. Horticulturae, 11(1), 62. [
DOI:10.3390/horticulturae11010062]
4. Albert, D., Nanaiah, K., & Sheshshayee, M. S. (2023). A Novel Automated Method for Screening Rice Genotypes for Root Growth Rate using ImageJ Software. Mysore Journal of Agricultural Sciences, 57(4), 89-97. [
DOI:10.23736/S1824-4777.23.01594-2]
5. Alboresi, A., Gestin, C., Leydecker, M.T., Bedu, M., Meyer, C. & Truong, H.N. (2005). Nitrate, a signal relieving seed dormancy in Arabidopsis. Plant, Cell & Environment, 28(4), 500-512. [
DOI:10.1111/j.1365-3040.2005.01292.x] [
PMID]
6. Al-Hadedy, S. H. A., Basheer, S. A., Idrees, M. S., & Al-Taee, K. A. Y. (2024). Sulfuric acid and hot water treatment effects on the seed germination and growth traits of Sesbania punicea L. SABRAO Journal of Breeding and Genetics, 56(1), 444-452. [
DOI:10.54910/sabrao2024.56.1.40]
7. Ali, F., Qanmber, G., Li, F., & Wang, Z. (2022). Updated role of ABA in seed maturation, dormancy, and germination. Journal of Advanced Research, 35, 199-214. [
DOI:10.1016/j.jare.2021.03.011] [
PMID] [
PMCID]
8. Anschütz, U., Becker, D., & Shabala, S. (2014). Going beyond nutrition: regulation of potassium homoeostasis as a common denominator of plant adaptive responses to environment. Journal of Plant Physiology, 171(9), 670-687. [
DOI:10.1016/j.jplph.2014.01.009] [
PMID]
9. Balouchi, H. R., & Modarres Sanavy, S. A. M. (2006). Effect of gibberellic acid, prechilling, sulfuric acid and potassium nitrate on seed germination and dormancy of annual medics. Pakistan Journal of Biological Sciences, 9(15), 2875-2880. [
DOI:10.3923/pjbs.2006.2875.2880]
10. Balouchi, H. R., Modarres Sanavy, S. A. M., & Alizadeh, B. (2008). Factors affecting dormancy and germination of two annual medic species. Iranian Journal of Biology, 21(2), 261-270. [In Persian]
11. Bani-Aameur, F., & Sipple-Michmerhuizen, J. (2001). Germination and seedling survival of Argan (Argania spinosa) under experimental saline conditions. Journal of Arid Environments, 49(3), 533-540. [
DOI:10.1006/jare.2001.0804]
12. Baskin, J. M., & Baskin, C. C. (2004). A classification system for seed dormancy. Seed Science Research, 14(1), 1-16. [
DOI:10.1079/SSR2003150] [
PMCID]
13. Bewley, J. D., Bradford, K. J., Hilhorst, H. W. M., & Nonogaki, H. (2013). Seeds: Physiology of development, germination and dormancy (3rd ed.). Springer. [
DOI:10.1007/978-1-4614-4693-4]
14. Bian, L., Yang, L., Wang, J. A., & Shen, H. L. (2013). Effects of KNO3 pretreatment and temperature on seed germination of Sorbus pohuashanensis. Journal of Forestry Research, 24(2), 309-316. [
DOI:10.1007/s11676-013-0354-9]
15. Black, M. & Bawley, J. D. (2000). Seed priming. Seed technology and its biological basis. Sheffield Academic Press, Sheffield, 287-325.
16. Boyle, T. H., & Hladun, K. (2005). Influence of seed size, testa color, scarification method, and immersion in cool or hot water on germination of Baptisia australis (L.) R. Br. seeds. Hortscience, 40(6), 1846-1849. [
DOI:10.21273/HORTSCI.40.6.1846]
17. Bush, E. W., Wilson, P., Shepard, D. P., & McClure, G. (2000). Enhancement of seed germination in common carpetgrass and centipedegrass seed. HortScience, 35(4), 769-770. [
DOI:10.21273/HORTSCI.35.4.769]
18. Cao, K., Mu, Y., Lu, S., & Zhao, Y. (2025). Regulatory mechanisms of exogenous Gibberellin on seed germination and transcriptomic responses in lomatogonium rotatum. Genes, 16(8), 878. [
DOI:10.3390/genes16080878] [
PMID] [
PMCID]
19. Cavallaro, V., Maucieri, C., Patanè, C., Fascella, G., Pellegrino, A., & Barbera, A. C. (2021). Polyphenols leaching and seed dormancy in carob (Ceratonia siliqua L.) in relation to hot water treatment. Acta Physiologiae Plantarum, 43(11), 141. [
DOI:10.1007/s11738-021-03308-z]
20. Devlin, P. F., Christie, J. M., & Terry, M. J. (2007). Many hands make light work. Journal of Experimental Botany, 58(12), 3071-3077. [
DOI:10.1093/jxb/erm251] [
PMID]
21. Duermeyer, L., Khodapanahi, E., Yan, D., Krapp, A., Rothstein, S. J., & Nambara, E. (2018). Regulation of seed dormancy and germination by nitrate. Seed Science Research, 28(3), 150-157.
https://doi.org/10.1017/S096025851800020X [
DOI:10.1017/s096025851800020x]
22. Fankhauser, C., & Chory, J. (1997). Light control of plant development. Annual Review of Cell and Developmental Biology, 13(1), 203-229. [
DOI:10.1146/annurev.cellbio.13.1.203] [
PMID]
23. Frett, J. J., Pill, W. G., & Morneau, D. C. (1991). A comparison of priming agents for tomato and asparagus seeds. HortScience, 26(9):1158-1159. https://doi.10.21273/HORTSCI.26.9.1158 [
DOI:10.21273/HORTSCI.26.9.1158]
24. Ghadiri, H., & Bagherani Torshiz, N. (2000). Effects of scarification and temperature on germination of licorice (Glycyrrhiza glabra L.) seeds. Journal of Agricultural Science and Technology, 2(4), 257-262. [In Persian]
25. Gurung, N., Swamy, G. S. K., Sarkar, S. K., & Ubale, N. B. (2014). Effect of chemicals and growth regulators on germination, vigour and growth of passion fruit (Passiflora edulis Sims.). The Bioscan, 9(1), 155-157.
26. Haj Sghaier, A., Tarnawa, Á., Khaeim, H., Kovács, G. P., Gyuricza, C., & Kende, Z. (2022). The effects of temperature and water on the seed germination and seedling development of rapeseed (Brassica napus L.). Plants, 11(21), 2819. [
DOI:10.3390/plants11212819] [
PMID] [
PMCID]
27. Han, Y., Hou, Z., Zhang, X., Yan, K., Liang, Z., & He, Q. (2022). Important changes in germination, seedling tolerance, and active components content due to drought stress on three licorice (Glycyrrhiza) species. Industrial Crops and Products, 175, 114240. [
DOI:10.1016/j.indcrop.2021.114240]
28. Harrison, R. J., Howieson, J. G., Yates, R. J., Norman, H. C., O'Hara, G. W., & Jaganathan, G. K. (2025). The impact of agricultural intensification on legume seeds with physical dormancy in sustainable farming systems. A review. Agronomy for Sustainable Development, 45(6), 64. [
DOI:10.1007/s13593-025-01058-7]
29. Hayashi, H., Hiraoka, N., Ikeshiro, Y., & Yamamoto, H. (1996). Organ specific localization of flavonoids in Glycyrrhiza glabra L. Plant Science, 116(2), 233-238. [
DOI:10.1016/0168-9452(96)04387-7]
30. He, J., Qin, L., Chong, E. L., Choong, T. W., & Lee, S. K. (2017). Plant growth and photosynthetic characteristics of Mesembryanthemum crystallinum grown aeroponically under different blue-and red-LEDs. Frontiers in Plant Science, 8, 361. [
DOI:10.3389/fpls.2017.00361] [
PMCID]
31. Hirsch, R. E., Lewis, B. D., Spalding, E. P., & Sussman, M. R. (1998). A role for the AKT1 potassium channel in plant nutrition. Science, 280(5365), 918-921. [
DOI:10.1126/science.280.5365.918] [
PMID]
32. Hosseini, M. S., Ebrahimi, M., Samsampour, D., Abadía, J., Khanahmadi, M., Amirian, R., & Gogorcena, Y. (2021). Association analysis and molecular tagging of phytochemicals in the endangered medicinal plant licorice (Glycyrrhiza glabra L.). Phytochemistry, 183, 112629. [
DOI:10.1016/j.phytochem.2020.112629] [
PMID]
33. Kafkafi, U., Xu, G., Imas, P., Magen, H., Tarchitzky, J., & Johnston, J. (2002). Potassium and chloride in crops and soils: The role of potassium chloride fertilizer in crop nutrition (IPI Research Topics No. 22). International Potash Institute.
34. Kaya, M., Kaya, G., Kaya, M. D., Atak, M., Saglam, S., Khawar, K. M., & Ciftci, C. Y. (2008). Interaction between seed size and NaCl on germination and early seedling growth of some Turkish cultivars of chickpea (Cicer arietinum L.). Journal of Zhejiang University Science, 9(5), 371-377. [
DOI:10.1631/jzus.B0720268] [
PMID] [
PMCID]
35. Khaeim, H., Kende, Z., Balla, I., Gyuricza, C., Eser, A., & Tarnawa, Á. (2022). The effect of temperature and water stresses on seed germination and seedling growth of wheat (Triticum aestivum L.). Sustainability, 14(7), 3887. [
DOI:10.3390/su14073887]
36. Kozhuharova, A., Nikolova, M., Stoyanov, S., Yankova-Tsvetkova, E., Ilinkin, V., Berkov, S., & Stanilova, M. (2024). Establishment of an ex situ collection of Glycyrrhiza glabra L. as a prerequisite for field cultivation in Bulgaria. Horticulturae, 10(1), 86. [
DOI:10.3390/horticulturae10010086]
37. Li, Y. J., Chen, J., Li, Y., Li, Q., Zheng, Y. F., Fu, Y., & Li, P. (2011). Screening and characterization of natural antioxidants in four Glycyrrhiza species by liquid chromatography coupled with electrospray ionization quadrupole time-of-flight tandem mass spectrometry. Journal of Chromatography, 1218(45), 8181-8191. [
DOI:10.1016/j.chroma.2011.09.030] [
PMID]
38. Madakadze, R., Chirco, E. M., & Khan, A. A. (1993). Seed germination of three flower species following matriconditioning under various environments. Journal of the American Society for Horticultural Science, 118(3), 330-334. [
DOI:10.21273/JASHS.118.3.330]
39. Mao, P.-s., Wang, Y.-h., Wang, X.-g., Lian, J.-j., & Huang, Y. (2008). Conditions and stimulation for germination in Glycyrrhiza uralensis fisch seeds. Agricultural Sciences in China, 7(12), 1438-1444. [
DOI:10.1016/S1671-2927(08)60400-9]
40. Moaaz Ali, M., Javed, T., Mauro, R. P., Shabbir, R., Afzal, I., & Yousef, A. F. (2020). Effect of seed priming with potassium nitrate on the performance of tomato. Agriculture, 10(11), 498. [
DOI:10.3390/agriculture10110498]
41. Moein, N., & Amooaghaie, R. (2025). The interactive effects of seed scarification and different nitrogenous compounds on the germination and seedling growth of Astragalus adscendens. Iranian Journal of Seed Research, 12(2), 133-156. [In Persian] [
DOI:10.66224/yujs.12.2.13]
42. Nasri, F., Koshesh Saba, M., Ghaderi, N., Mozafari, A. A., & Javadi, T. (2014). Improving germination and dormancy breaking in Alstroemeria ligtu hybrid seeds. Trakia Journal of Sciences, 12(1), 38-46.
43. Piri, R., Moradi, A. & Hosseini Moghaddam, M. (2018). Effect of accelerated aging and seed priming on germination and some biochemical indices of cumin (Cuminum Cyminum L.). Iranian Journal of Seed Sciences and Research, 5(1), 69-81. [In Persian] [
DOI:10.22124/jms.2018.2901]
44. Rafieiolhossaini, M., Tadayon, M. R., & Mazhari, M. (2015). The effect of dormancy breaking treatments on seed germination of Licorice medicinal plant. Journal of Crops Improvement, 16(4), 809-817. [In Persian] [
DOI:10.22059/jci.2015.53246]
45. Rasiukevičiūtė, N., Brazaitytė, A., Valiuškaitė, A., Vaštakaitė-Kairienė, V., Bylaitė, A., Duchovskis, P. & Samuolienė, G. (2020). Effect of different light-emitting diodes wavelengths on the growth of grey mold. Acta Horticulturae, 1296, 257-264. [
DOI:10.17660/ActaHortic.2020.1296.33]
46. Seng, M., & Cheong, E. J. (2020). Comparative study of various pretreatment on seed germination of Dalbergia cochinchinensis. Forest Science and Technology, 16(2), 68-74. [
DOI:10.1080/21580103.2020.1758801]
47. Sharifi-Rad, J., Quispe, C., Herrera-Bravo, J., Belén, L. H., Kaur, R., Kregiel, D., Uprety, Y., Beyatli, A., Yeskaliyeva, B., Kırkın, C., Özçelik, B., Sen, S., Acharya, K., Sharopov, F., Cruz-Martins, N., Kumar, M., Abdull Razis, A. F., Sunusi, U., Kamal, R. M., Shaheen, S., & Suleria, H. A. R. (2021). Glycyrrhiza genus: Enlightening phytochemical components for pharmacological and health-promoting abilities. Oxidative Medicine and Cellular Longevity, 2021, 7571132. [
DOI:10.1155/2021/7571132] [
PMID] [
PMCID]
48. Sheidai Karkaj, E., Amani, M., Younessi-Hamzekhanlu, M., & Jahantab, E. (2021). Effect of different treatments on the braking seed dormancy and germination indices in medical species of Glycyrrhiza glabra L. Journal of Plant Research (Iranian Journal of Biology), 34(4), 967-980. [In Persian]
49. Tapfumaneyi, L., Dube, P., Mavengahama, S., & Ngezimana, W. (2023). Effect of gibberellic acid and potassium nitrate seed treatments on the emergence and seedling vigor of Amaranth and Cleome gynandra. Agrosystems, Geosciences & Environment, 6, e20359. [
DOI:10.1002/agg2.20359]
50. Tehrani, P. F., Majd, A., Mahmoodzadeh, H., & Satari, T. N. (2016). Effect of red and blue light-emitting diodes on germination, morphological and anatomical features of Brassica napus. Advanced Studies in Biology, 8(4), 173-180. [
DOI:10.12988/asb.2016.6832]
51. Thongtip, A., Mosaleeyanon, K., Korinsak, S., Toojinda, T., Darwell, C. T., Chutimanukul, P., & Chutimanukul, P. (2022). Promotion of seed germination and early plant growth by KNO3 and light spectra in Ocimum tenuiflorum using a plant factory. Scientific Reports, 12, 6995. [
DOI:10.1038/s41598-022-11001-5] [
PMID] [
PMCID]
52. Toora, P. K., Tuan, P. A., Nguyen, T. N., Badea, A., & Ayele, B. T. (2024). Modulation in the ratio of abscisic acid to gibberellin level determines genetic variation of seed dormancy in barley (Hordeum vulgare L.). Journal of Plant Physiology, 301, 154301. [
DOI:10.1016/j.jplph.2024.154301] [
PMID]
53. Vance, C. P. (2001). Symbiotic nitrogen fixation and phosphorus acquisition. Plant nutrition in a world of declining renewable resources. Plant Physiology, 127(2), 390-397. [
DOI:10.1104/pp.010331] [
PMID] [
PMCID]
54. Villiers, T. A. (1975). Dormancy and the survival of plants, Edward Arnold publishers limited. Arnold, 71 p .
55. Wang, C., Chen, L., Xu, C., Shi, J., Chen, S., Tan, M., Chen, J., Zou, L., Chen, C., Liu, Z., & Liu, X. (2020). A comprehensive review for phytochemical, pharmacological, and biosynthesis studies on Glycyrrhiza spp. The American Journal of Chinese Medicine, 48(1), 17-45. [
DOI:10.1142/S0192415X20500020] [
PMID]
56. Wang, Y. R., Hanson, J., & Mariam, Y. W. (2011). Breaking hard seed dormancy in diverse accessions of five wild Vigna species by hot water and mechanical scarification. Seed Science and Technology, 39(1), 12-20. [
DOI:10.15258/sst.2011.39.1.02]
57. Wang, Y., & Wu, W. H. (2013). Potassium transport and signaling in higher plants. Annual Review of Plant Biology, 64, 451-476. [
DOI:10.1146/annurev-arplant-050312-120153] [
PMID]
58. Wei, Y., Wang, S., & Yu, D. (2023). The role of light quality in regulating early seedling development. Plants, 12(14), 2746. [
DOI:10.3390/plants12142746] [
PMID] [
PMCID]
59. Wen, Z., Lu, X., Wen, J., Wang, Z., & Chai, M. (2024). Physical seed dormancy in legumes: Molecular advances and perspectives. Plants, 13(11), 1473. [
DOI:10.3390/plants13111473] [
PMID] [
PMCID]
60. Yan, A., & Chen, Z. (2020). The control of seed dormancy and germination by temperature, light and nitrate. The Botanical Review, 86, 39-75. [
DOI:10.1007/s12229-020-09220-4]
61. Yan, B., Hou, J., Li, W., Luo, L., Ye, M., Zhao, Z., & Wang, W. (2023). A review on the plant resources of important medicinal licorice. Journal of Ethnopharmacology, 301, 115823. [
DOI:10.1016/j.jep.2022.115823] [
PMID]
62. Yang, R., Yuan, B. C., Ma, Y. S., Zhou, S., & Liu, Y. (2017). The anti-inflammatory activity of licorice, a widely used Chinese herb. Pharmaceutical Biology, 55(1), 5-18. [
DOI:10.1080/13880209.2016.1225775] [
PMID] [
PMCID]
63. Zhang, S., Hu, J., Zhang, Y., Xie, X. J., & Knapp, A. (2007). Seed priming with brassinolide improves lucerne (Medicago sativa L.) seed germination and seedling growth in relation to physiological changes under salinity stress. Australian Journal of Agricultural Research, 58(8), 811-815. [
DOI:10.1071/AR06253]