1. احمدی¬زاده اصفهانی، آ.، صدروی، م. و کاظمی، ش. (1398). تاثیر نانوکیتوزان بر بیماری سوختگی زودهنگام گوجهفرنگی. دانش بیماریشناسی گیاهی، 8(2)، 109-102.
2. اکبری کیارود، ل.، رهنما، ک.، گلمحمدی، م.، و نصراللهنژاد، س. (۱۴۰۰). بررسی اثر مهارکنندگی کیتوزان بر سیستم حدّنصاب احساس جمعیت باکتری Pseudomonas syringae pv. syringae 3289 . زیستشناسی میکروارگانیسمها، ۱۰(۳۷)، ۱۳-۲۴.
3. پورعزیز، پ.، و کولیوند، د. (۱۴۰۲). کاربرد پلیمرهای زیستی در مدیریت بیماریهای ویروسی گیاهان. دانش بیماریشناسی گیاهی، ۱۳(1)، ۱۲۵-۱۳۴.
4. مهدیان، ص. ع.، رمضانی دومیرکالیی، ا.، و تاجیک قنبری، م. ع. (۱۴۰۱). اثر اسانس مورد، کیتوزان و قارچکش تیابندازول بر کپک سبز مرکبات. دانش بیماریشناسی گیاهی، ۱۱(۱)، ۷۴-۸۸.
5. Akbari Kiarood, S. L., Rahnama, K., Golmohammadi, M., & Nasrolahnejad, S. (2020). Quorum-quenching endophytic bacteria inhibit disease caused by Pseudomonas syringae pv. syringae in Citrus cultivars. Journal of Basic Microbiology, 60(9), 746-757. [
DOI:10.1002/jobm.202000038]
6. Benhamou, N., & Thériault, G. (1992). Treatment with chitosan enhances resistance of tomato plants to the crown and root rot pathogen Fusarium oxysporum f. sp. radicis-lycopersici. Physiological and Molecular Plant Pathology, 41(1), 33-52.
https://doi.org/10.1016/0885-5765(92)90047-Y [
DOI:10.1016/0885-5765(92)90047-4]
7. Brulé, D., Héloir, M. C., Roudaire, T., Villette, J., Bonnet, S., Pascal, Y., Darblade, B., Crozier, P., Hugueney, P., Coma, V., & Poinssot, B. (2024). Increasing vineyard sustainability: Innovating a targeted chitosan-derived biocontrol solution to induce grapevine resistance against downy and powdery mildews. Frontiers in Plant Science, 15, 1360254. [
DOI:10.3389/fpls.2024.1360254]
8. Chandra, S., Chakraborty, N., Dasgupta, A., Sarkar, J., Panda, K., & Acharya, K. (2015). Chitosan nanoparticles: A positive modulator of innate immune responses in plants. Scientific Reports, 5(1), 15195. [
DOI:10.1038/srep15195]
9. Da Silva, J. T., Dantas de Sousa, P. H., Costa, A. F., De Menezes, L. B., Alves, S. F., Pellegrini, F., & Amaral, A. C. (2023). Fluconazole and propolis co-encapsulated in chitosan nanoparticles for the treatment of vulvovaginal candidiasis in a murine model. Medical Mycology, 61(11), myad113. [
DOI:10.1093/mmy/myad113]
10. Debnath, D., Samal, I., Mohapatra, C., Routray, S., Kesawat, M. S., & Labanya, R. (2022). Chitosan: An autocidal molecule of plant pathogenic fungus. Life, 12(11), 1908. [
DOI:10.3390/life12111908]
11. Dhillon, G. S., Kaur, S., Brar, S. K., & Verma, M. (2013). Green synthesis approach: Extraction of chitosan from fungus mycelia. Critical Reviews in Biotechnology, 33(4), 379-403. [
DOI:10.3109/07388551.2012.717217]
12. Dziedzic, I., & Kertmen, A. (2023). Methods of chitosan identification: History and trends. Letters in Applied NanoBioScience, 12(4), 94. [
DOI:10.33263/LIANBS124.094]
13. El Hadrami, A., Adam, L. R., El Hadrami, I., & Daayf, F. (2010). Chitosan in plant protection. Marine Drugs, 8(4), 968-987. [
DOI:10.3390/md8040968]
14. Hamza, Z. K., Abosereh, N. A., Salim, R. G., El-Sayed, A. F., & Aly, S. E. (2025). Unraveling the antifungal and aflatoxin B1 inhibitory efficacy of nano-encapsulated caraway essential oil based on molecular docking of major components. Scientific Reports, 15(1), 14951.
https://doi.org/10.1038/s41598-025-95557-y [
DOI:10.1038/s41598-025-92016-2]
15. Jabeen, N., & Ahmad, R. (2013). The activity of antioxidant enzymes in response to salt stress in safflower (Carthamus tinctorius L.) and sunflower (Helianthus annuus L.) seedlings raised from seed treated with chitosan. Journal of the Science of Food and Agriculture, 93(7), 1699-1705. [
DOI:10.1002/jsfa.5953]
16. Kappel, L., Kosa, N., & Gruber, S. (2022). The multilateral efficacy of chitosan and Trichoderma on sugar beet. Journal of Fungi, 8(2), 137. [
DOI:10.3390/jof8020137]
17. Kashyap, P. L., Xiang, X., & Heiden, P. (2015). Chitosan nanoparticle-based delivery systems for sustainable agriculture. International Journal of Biological Macromolecules, 77, 36-51. [
DOI:10.1016/j.ijbiomac.2015.02.039]
18. Liew, C. S., Lock, S. S. M., Mhd Yusof, S. M., Rawindran, H., Mong, G. R., Lim, J. W., Manogaran, M. D., & Suparmaniam, U. (2025). Chitin and chitosan production from Black Soldier Fly Larvae (Hermetia illucens) as bioresource: Current progress, applications, challenges and way forwards. Waste and Biomass Valorization.
https://doi.org/10.1007/s12649-025-03175-6 [
DOI:10.1007/s12649-025-02856-y]
19. Liu, T., Li, J., Tang, Q., Qiu, P., Gou, D., & Zhao, J. (2022). Chitosan-based materials: An overview of potential applications in food packaging. Foods, 11(10), 1490. [
DOI:10.3390/foods11101490]
20. Lopez-Nuñez, R., Prieto-Rubio, J., Bautista, I., Lidón-Cerezuela, A. L., Valverde-Urrea, M., Lopez-Moya, F., & Lopez-Llorca, L. V. (2025). Chitosan reduces naturally occurring plant pathogenic fungi and increases nematophagous fungus Purpureocillium in soil under field conditions. Frontiers in Agronomy, 6, 1502402. [
DOI:10.3389/fagro.2024.1502402]
21. Lopez-Nuñez, R., Suarez-Fernandez, M., Lopez-Moya, F., & Lopez-Llorca, L. V. (2022). Chitosan and nematophagous fungi for sustainable management of nematode pests. Frontiers in Fungal Biology, 3, 980341. [
DOI:10.3389/ffunb.2022.980341]
22. Malerba, M., & Cerana, R. (2016). Chitosan effects on plant systems. International Journal of Molecular Sciences, 17(7), 996. [
DOI:10.3390/ijms17070996]
23. Margaritopoulou, T., Sakellariou, A., Sofianos, G., Triviza, M. F., Stika, D. M., Tsiriva, D., Karaoglanidis, G., & Markellou, E. (2025). Chitosan nanoparticles loaded with jasmonic acid induce plants' resistance against Botrytis cinerea. Physiological and Molecular Plant Pathology, 135, 102887. [
DOI:10.1016/j.pmpp.2025.102887]
24. Pellis, A., Guebitz, G. M., & Nyanhongo, G. S. (2022). Chitosan: Sources, processing and modification techniques. Gels, 8(7), 393. [
DOI:10.3390/gels8070393]
25. Raghavan, N., & Pathan, E. K. (2025). A comprehensive account of fungal chitin deacetylases: Aspects and prospects. International Journal of Biological Macromolecules, 267(Part 1), 142705.
https://doi.org/10.1016/j.ijbiomac.2025.142705 [
DOI:10.1016/j.ijbiomac.2024.142705]
26. Reshad, R. A. I., Jishan, T. A., & Chowdhury, N. N. (2021). Chitosan and its broad applications: A brief review. SSRN Electronic Journal. [
DOI:10.2139/ssrn.3842055]
27. Rojas-Pirela, M., Carillo, P., Lárez-Velásquez, C., & Romanazzi, G. (2024). Effects of chitosan on plant growth under stress conditions: Similarities with plant growth promoting bacteria. Frontiers in Plant Science, 15, 1423949. [
DOI:10.3389/fpls.2024.1423949]
28. Samarah, N. H., Al-Quraan, N. A., Massad, R. S., & Welbaum, G. E. (2020). Treatment of bell pepper (Capsicum annuum L.) seeds with chitosan increases chitinase and glucanase activities and enhances emergence in a standard cold test. Scientia Horticulturae, 269, 109393. [
DOI:10.1016/j.scienta.2020.109393]
29. Shahifar, A., Ghasemi, Z., & Shahifar, R. (2024). Extraction of chitosan from the Penaeus vannamei shrimp shell and investigation of its effects on quality preservation of grape and strawberry fruits during storage. Fisheries Science and Technology, 13(1), 535-547. [In Persian].
30. Sharif, R., Mujtaba, M., Ur Rahman, M., Shalmani, A., Ahmad, H., Anwar, T., Tianchan, D., & Wang, X. (2018). The multifunctional role of chitosan in horticultural crops: A review. Molecules, 23(4), 872. [
DOI:10.3390/molecules23040872]
31. Shcherban, A. B. (2023). Chitosan and its derivatives as promising plant protection tools. Vavilov Journal of Genetics and Breeding, 27(8), 1010-1018. [
DOI:10.18699/VJGB-23-118]
32. Shetranjiwalla, S., & Ononiwu, A. (2025). Identifying barriers to scaled-up production and commercialization of chitin and chitosan using green technologies: A review and quantitative green chemistry assessment. International Journal of Biological Macromolecules, 305(Part 1), 132456. [
DOI:10.1016/j.ijbiomac.2024.132456]
33. Suwançaikasem, P., Idnurm, A., Selby-Pham, J., Walker, R., & Boughton, B. A. (2024). The impacts of chitosan on plant root systems and its potential to be used for controlling fungal diseases in agriculture. Journal of Plant Growth Regulation, 43(5), 1234-1248.
https://doi.org/10.1007/s00344-024-11356-1 [
DOI:10.1007/s00344-024-11462-2]
34. Varun, T. K., Senani, S., Jayapal, N., Chikkerur, J., Roy, S., Tekulapally, V. B., Gautam, M., & Kumar, N. (2017). Extraction of chitosan and its oligomers from shrimp shell waste, their characterization and antimicrobial effect. Veterinary World, 10(2), 170-175. [
DOI:10.14202/vetworld.2017.170-175]
35. Younes, I., & Rinaudo, M. (2015). Chitin and chitosan preparation from marine sources: Structure, properties and applications. Marine Drugs, 13(3), 1133-1174. [
DOI:10.3390/md13031133]
36. Zhang, L., Yang, Y., Zhu, Y., Hu, H., Jia, Q., Sun, C., Zhu, X., & Liu, Y. (2024). Antifungal activity and mechanism of chitosan against Fusarium solani causing ginger soft rot during postharvest storage. Postharvest Biology and Technology, 208, 112680. [
DOI:10.1016/j.postharvbio.2023.112680]
37. Zhang, Z., Ma, Z., Song, L., & Farag, M. A. (2024). Maximizing crustaceans (shrimp, crab and lobster) by-products value for optimum valorization practices: A comparative review of their active ingredients extraction, bioprocesses and applications. Journal of Advanced Research, 57, 59-76. [
DOI:10.1016/j.jare.2023.04.019]